Squalius
| Squalius | |
|---|---|
 | |
| Squalius carolitertii | |
 | |
| Squalius pyrenaicus | |
| Scientific classification | |
| Kingdom: | Animalia |
| Phylum: | Chordata |
| Class: | Actinopterygii |
| Order: | Cypriniformes |
| Family: | Cyprinidae |
| Subfamily: | Leuciscinae |
| Genus: | Squalius Bonaparte, 1837 |
| Type species | |
| Leuciscus squalus Bonaparte, 1837 | |
Squalius is a genus of fish in the family Cyprinidae, commonly known as chubs. This genus belongs to the subfamiliy Leuciscinae. The Eurasian daces, Leuciscus are close relatives. The present genus was formerly entirely included there and their delimitation with respect to each other is somewhat uncertain. A number of species have in recent times been moved from one genus to the other. Generally, Leuciscus in the modern sense is widespread across Eurasia, whereas Squalius is almost limited to Europe proper.[1]
Hybridization is not rare in the Cyprinidae, including this genus. S. alburnoides is known to be of ancient hybrid origin, with the paternal lineage deriving from a prehistoric species related to Anaecypris; the latter mated with ancestral S. pyrenaicus. Present-day S. alburnoides mates with sympatric congeners of other species.[2]
Species
There are currently 49 recognized species in this genus:
- Squalius adanaensis Turan, Kottelat & Doğan, 2013 (Adana chub) [3]
- Squalius agdamicus S. N. Kamensky, 1901
- Squalius alburnoides (Steindachner, 1866) [2]
- Squalius anatolicus (Bogutskaya, 1997) (Beyşehir chub) [4]
- Squalius aphipsi (A. I. Aleksandrov, 1927) (Aphips chub)
- Squalius aradensis (M. M. Coelho, Bogutskaya, Rodrigues & Collares-Pereira, 1998)
- Squalius aristotelis Özuluğ & Freyhof, 2011 (Tuzla chub) [4]
- Squalius berak Heckel, 1843 (Mesopotamian chub) [3]
- Squalius cappadocicus Özuluğ & Freyhof, 2011 (Cappadocian chub) [4]
- Squalius carinus Özuluğ & Freyhof, 2011 (Chocolate chub) [4]
- Squalius carolitertii (Doadrio, 1988)
- Squalius castellanus Doadrio, Perea & F. Alonso, 2007
- Squalius cephaloides (Battalgil, 1942) (Thick-lipped chub) [4]
- Squalius cephalus (Linnaeus, 1758) (European chub)
- Squalius cii (J. Richardson, 1857) [4]
- Squalius fellowesii (Günther, 1868) (Aegean chub) [4]
- Squalius ghigii (Gianferrari, 1927) (Rhodes chub)
- Squalius illyricus Heckel & Kner, 1857
- Squalius irideus (Ladiges, 1960)
- Squalius janae Bogutskaya & Zupančič, 2010
- Squalius keadicus (Stephanidis, 1971)
- Squalius kosswigi (M. S. Karaman (sr), 1972) [4]
- Squalius kottelati Turan, Yilmaz & Kaya, 2009 (Striped chub)
- Squalius laietanus Doadrio, Kottelat & de Sostoa, 2007
- Squalius lepidus Heckel, 1843
- Squalius lucumonis (Bianco, 1983)
- Squalius malacitanus Doadrio & Carmona, 2006
- Squalius microlepis Heckel, 1843
- Squalius moreoticus (Stephanidis, 1971) (Stymphalia chub)
- Squalius namak Khaefi, Esmaeili, Sayyadzadeh, Geiger & Freyhof, 2016 [5]
- Squalius orientalis (Nordmann, 1840)
- Squalius orpheus Kottelat & Economidis, 2006 (Orpheus chub)
- Squalius pamvoticus (Stephanidis, 1939) (Pamvotis chub)
- Squalius peloponensis (Valenciennes, 1844) (Peloponnese chub)
- Squalius platyceps Zupančič, Marić, Naseka & Bogutskaya, 2010
- Squalius prespensis (Fowler, 1977)
- Squalius pursakensis (Hankó, 1925) (Sakarya chub) [4]
- Squalius pyrenaicus (Günther, 1868)
- Squalius recurvirostris Özuluğ & Freyhof, 2011 (Akşehir chub) [4]
- Squalius seyhanensis Turan, Kottelat & Doğan, 2013 (Seyhan chub) [3]
- Squalius spurius Heckel, 1843 (Orontes chub)
- Squalius squalus (Bonaparte, 1837)
- Squalius svallize Heckel & Kner, 1857
- Squalius tenellus Heckel, 1843
- Squalius torgalensis (M. M. Coelho, Bogutskaya, Rodrigues & Collares-Pereira, 1998)
- Squalius turcicus De Filippi, 1865 (Transcaucasian chub)
- Squalius valentinus Doadrio & Carmona, 2006
- Squalius vardarensis S. L. Karaman, 1928
- Squalius zrmanjae S. L. Karaman, 1928
References
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- ↑ Froese, Rainer, and Daniel Pauly, eds. (2016). Species of Squalius in FishBase. June 2016 version.
- 1 2 Collares-Pereira, M.J. & Coelho, M.M. (2010): Reconfirming the hybrid origin and generic status of the Iberian cyprinid complex Squalius alburnoides. Journal of Fish Biology, 76 (3): 707–715.
- 1 2 3 Turan, D., Kottelat, M. & Doğan, E. (2013): Two new species of Squalius, S. adanaensis and S. seyhanensis (Teleostei: Cyprinidae), from the Seyhan River in Turkey. Zootaxa, 3637 (3): 308–324.
- 1 2 3 4 5 6 7 8 9 10 Özuluğ, M. & Freyhof, J. (2011): Revision of the genus Squalius in Western and Central Anatolia, with description of four new species (Teleostei: Cyprinidae). Ichthyological Exploration of Freshwaters, 22 (2): 107-148.
- ↑ Khaefi, R., Esmaeili, H.R., Sayyadzadeh, G., Geiger, M.F. & Freyhof, J. (2016): Squalius namak, a new chub from Lake Namak basin in Iran (Teleostei: Cyprinidae). Zootaxa, 4169 (1): 145–159.
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